The San Francisco 49ers will have their entire defense returning next season, but the team made several moves Thursday to give it a different look in 2012.
The 49ers released cornerback Shawntae Spencer on Thursday after he spent eight seasons with the team.
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The 25-year-old Manningham had 39 catches for 523 yards and four touchdowns last season, his fourth in the NFL, while playing behind Hakeem Nicks and Victor Cruz. Lloyd, who played three seasons for San Francisco from 2003-05, made 70 receptions for 966 yards and five touchdowns in 11 games for St. Louis last season.
The Associated Press contributed to this report.